Eastern Michigan University
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Dr. Steven Francoeur


Steven Francoeur 401F Mark Jefferson Science Complex




B.Sc., Bowling Green State University, 1994

M.Sc., Bowling Green State University, 1997

Ph.D., University of Alabama, 2001

Interests and Expertise

Virtually all submerged surfaces in lakes, rivers, and wetlands are covered with microbes. These surface-associated microbial communities are composed of algae, bacteria, fungi, and protists, and are known as periphyton. My research is mainly focused on the ecology of periphyton, with emphasis on the algal component. The two main themes of my research are: 1) understanding how biological, physical, and chemical factors affect the abundance, distribution, and activity of periphytic microbes, and 2) understanding microbial interactions within periphyton communities. Examples include quantifying the effects of light and nutrients on benthic algae in the Great Lakes, and investigating the influence of algal photosynthesis on bacteria and fungi within periphyton communities.


Kuehn, K. A, S. N. Francoeur, R. H. Findlay and R. K. Neely. 2014. Priming in the microbial landscape: Periphytic algal stimulation of litter-associated microbial decomposers. Ecology 95:749-762.
Francoeur, S. N., K. A. Peters-Winslow, D. Miller, C. A. Stow, Y-K. Cha, and S. D. Peacor. 2014. Spatial and temporal patterns of macroscopic benthic primary producers in Saginaw Bay, Lake Huron. Journal of Great Lakes Research 40(Supplement):53-63.
Peters-Winslow, K. A., S. N. Francoeur, and S. D. Peacor. 2014. The influence of light and nutrients on benthic filamentous algal growth: A case study of Saginaw Bay, Lake Huron. Journal of Great Lakes Research 40(Supplement):64-74.
Stow, C. A., J. Dyble, D. R. Kashian, T. H. Johengen, K. Peters-Winslow, S. D. Peacor, S. N. Francoeur, A. M. Burtner, D. Palladino, N. Morehead, D. Gossiaux,  Y-K. Cha, S. S. Qian, and
D. Miller. 2014. Phosphorus targets and eutrophication objectives in Saginaw Bay: A 35 year assessment. Journal of Great Lakes Research 40(Supplement):4-10.
Francoeur, S. N., S. T. Rier, and S. B. Whorley. 2013. Methods for sampling and analyzing wetland algae. Chapter 1 in Wetland Techniques, Vol. 2 (J. T. Anderson, W. C. Conway, and C. A. Davis, eds.). Springer, New York, NY.
Whorley, S. B., and S. N. Francoeur. 2013. Active fluorimetry improves nutrient-diffusing substrata bioassay. Freshwater Science 32:108-115.
Cook, L. J., and S. N. Francoeur. 2013. Effects of simulated short-term road salt exposure on lotic periphyton function. Journal of Freshwater Ecology 28:211-223.
Kuehn, K. A., B. M. Ohsowski, S. N. Francoeur, and R. K. Neely. 2011. Contributions of fungi to carbon flow and nutrient cycling from standing-dead Typha angustifolia leaf litter in a temperate freshwater marsh. Limnology and Oceanography 56:529-539.
Hill, W. R., B. J. Roberts, S. N. Francoeur, and S. E. Fanta. 2011. Resource synergy and the autotrophic:heterotrophic balance in experimental streams. Journal of Ecology 99:454-463.
West, B. M., P. Liggit, D. L. Clemans, and S. N. Francoeur. 2011. Antibiotic resistance, gene transfer, and water quality patterns observed in waterways near CAFO farms and wastewater treatment facilities. Water Air and Soil Pollution 217:473-489.
Liess, A., and S. N. Francoeur. 2011. Substratum-associated microbiota. Water Environment Research Literature Review 83:1704-1732.

Courses Taught

BIO 110 Introductory Biology I

BIO 310 Ecology

BIO 410 Limnology

BIO 456 Freshwater Algae

BIO 509 Statistics for Biologists

BIO 510 Limnology

BIO 556 Freshwater Algae

BIO 413 Wetland Ecosystems

BIO 513 Wetland Ecosystems

CSIE 150L Cyanobacteria in the Huron River

The Biology Department is part of the College of Arts & Sciences, 214 Pray-Harrold