Eastern Michigan University
direct edit

Dr. Robert S. Winning


Robert Winning 401D Mark Jefferson Science Complex




B.Sc., University of Calgary, 1983

M.Sc., University of Calgary, 1986

Ph.D., University of Waterloo, 1990

Interests and Expertise

My general research interests concern the molecular analysis of early developmental events in amphibian embryogenesis. More specifically, I am interested in how cellular interactions and cell signaling affect cell fate and cell behavior in embryos. Current research involves examination of the function of a family of receptor tyrosine kinases in embryos of the frog Xenopus laevis.


Winning, R.S. and Krull, C.E., 2011. Knockdown of Ephrin-A5 Expression by 40% Does not Affect Motor Axon Growth or Migration into the Chick Hindlimb. Int. J. Mol. Sci. 12: 8362-8371.

Winning, R. S., Ward, E. K., Scales, J. B., and Walker, G. K., 2002. EphA4 catalytic activity causes inhibition of RhoA GTPase in Xenopus laevis embryos. Differentiation 70: 46-55.

Winning, R. S., Wyman, T. L., and Walker, G. K., 2001. EphA4 activity causes cell shape change and a loss of cell polarity in Xenopus laevis embryos. Differentiation 68: 126-132.

Winning, R.S., Scales, J.B., and Sargent, T.D., 1996. Disruption of cell adhesion in Xenopus embryos by Pagliaccio, an Eph- class receptor tyrosine kinase. Developmental Biology 179: 309-319.

Scales, J.B., Winning, R.S., Renaud, C.S., Shea, L.J., and Sargent, T.D., 1995. Novel members of the Eph receptor tyrosine kinase subfamily expressed during Xenopus development. Oncogene 11: 1745-1752.

Courses Taught

BIO 110 Introductory Biology I

BIO 257 Human Embryonic Development

BIO 301 Genetics

BIO 443 Developmental Biology

BIO 540 Cell Biology

BIO 543 Developmental Biology


Recipient of the 2010 Ron Collins Distinguished Faculty Award Winner for Teaching

Michigan Distinguished Professor of the Year, 2011

The Biology Department is part of the College of Arts & Sciences, 214 Pray-Harrold