Robert S. Winning

A photo of Robert Winning



401D Science Complex


[email protected]


  • BS, University of Calgary, 1983
  • MS, University of Calgary, 1986
  • Ph.D., University of Waterloo, 1990

Interests and Expertise

My general research interests concern the molecular analysis of early developmental events in amphibian embryogenesis. More specifically, I am interested in how cellular interactions and cell signaling affect cell fate and cell behavior in embryos. Current research involves examination of the function of a family of receptor tyrosine kinases in embryos of the frog Xenopus laevis.


  • BIO 110 Introductory Biology I
  • BIO 257 Human Embryonic Development
  • BIO 301 Genetics
  • BIO 443 Developmental Biology
  • BIO 540 Cell Biology
  • BIO 543 Developmental Biology


  • Recipient of the 2010 Ron Collins Distinguished Faculty Award Winner for Teaching
  • Michigan Distinguished Professor of the Year, 2011

Publications and Presentations

  • Winning, R.S. and Krull, C.E., 2011. Knockdown of Ephrin-A5 Expression by 40% Does not Affect Motor Axon Growth or Migration into the Chick Hindlimb. Int. J. Mol. Sci. 12: 8362–8371.
  • Winning, R. S., Ward, E. K., Scales, J. B., and Walker, G. K., 2002. EphA4 catalytic activity causes inhibition of RhoA GTPase in Xenopus laevis embryos. Differentiation 70: 4655.
  • Winning, R. S., Wyman, T. L., and Walker, G. K., 2001. EphA4 activity causes cell shape change and a loss of cell polarity in Xenopus laevis embryos. Differentiation 68: 126132.
  • Winning, R.S., Scales, J.B., and Sargent, T.D., 1996. Disruption of cell adhesion in Xenopus embryos by Pagliaccio, an Eph-class receptor tyrosine kinase. Developmental Biology 179: 309319.
  • Scales, J.B., Winning, R.S., Renaud, C.S., Shea, L.J., and Sargent, T.D., 1995. Novel members of the Eph receptor tyrosine kinase subfamily expressed during Xenopus development. Oncogene 11: 17451752.